Analysis and evaluation of phenotypic character diversity of olive germplasm resources

doi:10.3969/j.issn.1004-1524.2023.05.04

Acta Agriculturae Zhejiangensis ›› 2023, Vol. 35 ›› Issue (5): 1001-1015.DOI: 10.3969/j.issn.1004-1524.2023.05.04

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Analysis and evaluation of phenotypic character diversity of olive germplasm resources

QI Jianli¹^,²(), ZHANG Rong¹^,², WU Wenjun¹^,², JIANG Chengying¹^,²^,^*(), ZHAO Mengjiong¹^,², JIN Gaoming¹^,², CHEN Weiqing¹^,²

1. Center for Olive Research and Technology, Gansu Academy of Forestry Sciences, Lanzhou 730020, China
2. National Forestry Grassland Olive Engineering Technology Research Center, Lanzhou 730020, China

Received:2022-08-31 Online:2023-05-25 Published:2023-06-01

Abstract

Abstract:

In order to explore the genetic diversity and variation characteristics of phenotypic traits of olive germplasm resources in Gansu Province, and to provide a theoretical reference for the selection, evaluation, development and utilization of local olive germplasm resources, 11 characters of 35 olive germplasm resources were described, and 16 phenotypic characters were analyzed by diversity, correlation, principal component analysis and cluster analysis. The results showed that there were significant differences among the phenotypic characters of different olive germplasm resources, and the coefficient of variation ranged from 8.64% to 55.87%. The variation coefficient of single fruit weight was the largest, and the variation coefficient of flesh rate was the smallest. Correlation analysis showed that fruit longitudinal diameter, fruit transverse diameter, single fruit weight, pulp thickness, fruit core longitudinal diameter, fruit core transverse diameter, fruit core weight and pulp rate were significantly (P <0.01) positively correlated with each other. Principal component analysis showed that 16 phenotypic traits could be extracted into 5 principal components, with a cumulative contribution rate of 91.633%. Cluster analysis showed that 35 olive germplasm resources could be divided into 5 categories at Euclidean distances of 10, which could be used as 5 different breeding directions. Among them, the highest oil content of dried and fresh fruits of Chemlal de kabylie could be grouped into one category, which could be used as special germplasm for mining; M1 had the least water content, and the oil content of fresh and dry fruits was only lower than that of Chemlal de kabylie, which was divided into oil products; Berat and LYuan Manzanilla had the highest water content and low oil content in fresh fruit, which could be divided into fruit products; The other two types of germplasm could be excavated and utilized in combination with other physiological indicators. This study showed that the phenotypic traits of different olive germplasm resources were significantly different, and the biodiversity was rich. The phenotypic traits of Nikiskii, LYuan Manzanilla, M1, Chemlal de kabylie and other germplasm were prominent, which could be used as parents for selection. Through correlation analysis, 7 indexes, such as single fruit weight, fruit shape index, stone index, fresh fruit oil rate and dry fruit oil rate, were selected as important indexes for selection, breeding and identification of olive germplasm resources.

Key words: olive, germplasm resource, phenotypic traits, diversity

CLC Number:

S565.7

QI Jianli, ZHANG Rong, WU Wenjun, JIANG Chengying, ZHAO Mengjiong, JIN Gaoming, CHEN Weiqing. Analysis and evaluation of phenotypic character diversity of olive germplasm resources[J]. Acta Agriculturae Zhejiangensis, 2023, 35(5): 1001-1015.

Figures/Tables 9

References 24

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序号 No.	种质 Germplasm	类型 Type	来源 Origin	原产国 Country of origin	序号 No.	种质 Germplasm	类型 Type	来源 Origin	原产国 Country of origin
1	尼2 N2	引进品种 Introduced variety	陇南 Longnan	阿普歇伦半岛 Apsheron	19	拉多利亚2 Ladolia2	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
2	贝拉 Berat	引进品种 Introduced variety	江苏 Jiangsu	阿尔巴尼亚 Albania	20	岚桂塔 Blanqueta	引进品种 Introduced variety	四川 Sichuan	西班牙 Spain
3	贺吉布兰卡 Hojiblanca	引进品种 Introduced variety	陇南 Longnan	西班牙 Spain	21	科拉蒂 Coratina	引进品种 Introduced variety	陇南 Longnan	意大利 Italy
4	M1	选育品种 Breeding variety	云南 Yunnan	中国China	22	希腊-3 Greece-3	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
5	M2	选育品种 Breeding variety	云南 Yunnan	中国China	23	绿源小苹果 LYuan Manzanilla	选育品种 Breeding variety	云南 Yunnan	中国China
6	M4	选育品种 Breeding variety	云南 Yunnan	中国China	24	卡林 Kaliniot	引进品种 Introduced variety	云南 Yunnan	阿尔巴尼亚 Albania
7	西格伊司27 Seeger Ismail27	引进品种 Introduced variety	四川 Sichuan	阿尔及利亚 Algeria	25	绿源1 LYuan-1	选育品种 Breeding variety	云南 Yunnan	中国China
8	莱星 Leccino	引进品种 Introduced variety	陇南 Longnan	意大利 Italy	26	鄂植8 EZ-8	选育品种 Breeding variety	陇南 Longnan	湖北 Hubei
9	云杂1 YunZ-1	选育品种 Breeding variety	云南 Yunnan		27	皮瓜儿 Picual	引进品种 Introduced variety	陇南 Longnan	西班牙 Spain
10	云杂2 YunZ-2	选育品种 Breeding variety	云南 Yunnan	中国China	28	米提尼 Adramittini	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
11	云杂3 YunZ-3	选育品种 Breeding variety	云南 Yunnan	中国China	29	小尼 Nikiskii	引进品种 Introduced variety	陇南 Longnan	阿普歇伦半岛 Apsheron
12	边克丽娜 Biancolilla	引进品种 Imported variety	陇南 Longnan	意大利 Italy	30	会理-X Huili-X	引进品种 Introduced variety	四川 Sichuan	中国China
13	软阿斯 Ascolano Tenera	引进品种 Introduced variety	陇南 Longnan	意大利 Italy	31	城固32 Chenggu-32	选育品种 Breeding variety	陇南 Longnan	中国China
14	豆果 Arbequina	引进品种 Introduced variety	湖北 Hubei	西班牙 Spain	32	奇迹 Koroneiki	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
15	切姆拉里 Chemlal de kabylie	引进品种 Introduced variety	湖北 Hubei	阿尔及利亚 Algeria	33	白橄榄 Ulliri Bardhe	引进品种 Introduced variety	陇南 Longnan	阿尔巴尼亚 Albania
16	苏里 Soury	引进品种 Introduced variety	云南 Yunnan	黎巴嫩 Lebanon	34	恩帕特雷 Empeltre	引进品种 Introduced variety	陇南 Longnan	西班牙 Spain
17	马托迪 Mastoidis	引进品种 Introduced variety	湖北 Hubei	希腊 Greece	35	西阿基 Thiaki	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
18	爱桑 Elbasan	引进品种 Introduced variety	云南 Yunnan	阿尔巴尼亚 Albania

序号 No.	种质 Germplasm	类型 Type	来源 Origin	原产国 Country of origin	序号 No.	种质 Germplasm	类型 Type	来源 Origin	原产国 Country of origin
1	尼2 N2	引进品种 Introduced variety	陇南 Longnan	阿普歇伦半岛 Apsheron	19	拉多利亚2 Ladolia2	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
2	贝拉 Berat	引进品种 Introduced variety	江苏 Jiangsu	阿尔巴尼亚 Albania	20	岚桂塔 Blanqueta	引进品种 Introduced variety	四川 Sichuan	西班牙 Spain
3	贺吉布兰卡 Hojiblanca	引进品种 Introduced variety	陇南 Longnan	西班牙 Spain	21	科拉蒂 Coratina	引进品种 Introduced variety	陇南 Longnan	意大利 Italy
4	M1	选育品种 Breeding variety	云南 Yunnan	中国China	22	希腊-3 Greece-3	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
5	M2	选育品种 Breeding variety	云南 Yunnan	中国China	23	绿源小苹果 LYuan Manzanilla	选育品种 Breeding variety	云南 Yunnan	中国China
6	M4	选育品种 Breeding variety	云南 Yunnan	中国China	24	卡林 Kaliniot	引进品种 Introduced variety	云南 Yunnan	阿尔巴尼亚 Albania
7	西格伊司27 Seeger Ismail27	引进品种 Introduced variety	四川 Sichuan	阿尔及利亚 Algeria	25	绿源1 LYuan-1	选育品种 Breeding variety	云南 Yunnan	中国China
8	莱星 Leccino	引进品种 Introduced variety	陇南 Longnan	意大利 Italy	26	鄂植8 EZ-8	选育品种 Breeding variety	陇南 Longnan	湖北 Hubei
9	云杂1 YunZ-1	选育品种 Breeding variety	云南 Yunnan		27	皮瓜儿 Picual	引进品种 Introduced variety	陇南 Longnan	西班牙 Spain
10	云杂2 YunZ-2	选育品种 Breeding variety	云南 Yunnan	中国China	28	米提尼 Adramittini	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
11	云杂3 YunZ-3	选育品种 Breeding variety	云南 Yunnan	中国China	29	小尼 Nikiskii	引进品种 Introduced variety	陇南 Longnan	阿普歇伦半岛 Apsheron
12	边克丽娜 Biancolilla	引进品种 Imported variety	陇南 Longnan	意大利 Italy	30	会理-X Huili-X	引进品种 Introduced variety	四川 Sichuan	中国China
13	软阿斯 Ascolano Tenera	引进品种 Introduced variety	陇南 Longnan	意大利 Italy	31	城固32 Chenggu-32	选育品种 Breeding variety	陇南 Longnan	中国China
14	豆果 Arbequina	引进品种 Introduced variety	湖北 Hubei	西班牙 Spain	32	奇迹 Koroneiki	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
15	切姆拉里 Chemlal de kabylie	引进品种 Introduced variety	湖北 Hubei	阿尔及利亚 Algeria	33	白橄榄 Ulliri Bardhe	引进品种 Introduced variety	陇南 Longnan	阿尔巴尼亚 Albania
16	苏里 Soury	引进品种 Introduced variety	云南 Yunnan	黎巴嫩 Lebanon	34	恩帕特雷 Empeltre	引进品种 Introduced variety	陇南 Longnan	西班牙 Spain
17	马托迪 Mastoidis	引进品种 Introduced variety	湖北 Hubei	希腊 Greece	35	西阿基 Thiaki	引进品种 Introduced variety	陇南 Longnan	希腊 Greece
18	爱桑 Elbasan	引进品种 Introduced variety	云南 Yunnan	阿尔巴尼亚 Albania

序号 No.	形状 Shape	对称性 Symmetry	果相 Apex	果基 Base	果顶凸起 Nipple	果粉 Fruit powder	果面斑点 Presence of lenticel
1	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	少Few
2	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	多Many
3	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	多Many
4	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	中Medium
5	卵形Ovoid	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	中Medium
6	卵形Ovoid	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	多Many
7	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	多Many
8	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
9	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
10	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
11	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	中Medium
12	椭圆形Elliptic	稍不对称Slightly asymmetric	尖Pointed	截尾Truncation	有Yes	有Yes	中Medium
13	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	多Many
14	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	截尾Truncation	无None	有Yes	少Few
15	椭圆形Elliptic	稍不对称Slightly asymmetric	尖Pointed	圆Round	有Yes	有Yes	少Few
16	卵形Ovoid	稍不对称Slightly asymmetric	尖Pointed	截尾Truncation	有Yes	有Yes	中Medium
17	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	多Many
18	卵形Ovoid	稍不对称Slightly asymmetric	尖Pointed	截尾Truncation	有Yes	有Yes	少Few
19	椭圆形Elliptic	不对称Asymmetry	尖Pointed	圆Round	有Yes	有Yes	少Few
20	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
21	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	无None	多Many
22	长椭圆形Oblong	稍不对称Slightly asymmetric	尖Pointed	圆Round	有Yes	有Yes	少Few
23	卵形Ovoid	完全对称Complete symmetry	尖Pointed	截尾Truncation	有Yes	有Yes	少Few
24	细长形Elongate	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	多Many
25	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	多Many
26	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
27	卵形Ovoid	不对称Asymmetry	尖Pointed	圆Round	有Yes	有Yes	多Many
28	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	多Many
29	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	中Medium
30	卵形Ovoid	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	多Many
31	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
32	卵形Ovoid	稍不对称Slightly asymmetric	尖Pointed	圆Round	有Yes	有Yes	少Few
33	圆球形Spheroid	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	多Many
34	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	少Few
35	卵形Ovoid	不对称Asymmetry	尖Pointed	圆Round	有Yes	有Yes	中Medium

序号 No.	形状 Shape	对称性 Symmetry	果相 Apex	果基 Base	果顶凸起 Nipple	果粉 Fruit powder	果面斑点 Presence of lenticel
1	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	少Few
2	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	多Many
3	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	多Many
4	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	中Medium
5	卵形Ovoid	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	中Medium
6	卵形Ovoid	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	多Many
7	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	多Many
8	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
9	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
10	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
11	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	中Medium
12	椭圆形Elliptic	稍不对称Slightly asymmetric	尖Pointed	截尾Truncation	有Yes	有Yes	中Medium
13	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	多Many
14	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	截尾Truncation	无None	有Yes	少Few
15	椭圆形Elliptic	稍不对称Slightly asymmetric	尖Pointed	圆Round	有Yes	有Yes	少Few
16	卵形Ovoid	稍不对称Slightly asymmetric	尖Pointed	截尾Truncation	有Yes	有Yes	中Medium
17	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	多Many
18	卵形Ovoid	稍不对称Slightly asymmetric	尖Pointed	截尾Truncation	有Yes	有Yes	少Few
19	椭圆形Elliptic	不对称Asymmetry	尖Pointed	圆Round	有Yes	有Yes	少Few
20	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
21	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	无None	多Many
22	长椭圆形Oblong	稍不对称Slightly asymmetric	尖Pointed	圆Round	有Yes	有Yes	少Few
23	卵形Ovoid	完全对称Complete symmetry	尖Pointed	截尾Truncation	有Yes	有Yes	少Few
24	细长形Elongate	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	多Many
25	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	多Many
26	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
27	卵形Ovoid	不对称Asymmetry	尖Pointed	圆Round	有Yes	有Yes	多Many
28	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	多Many
29	椭圆形Elliptic	完全对称Complete symmetry	圆Round	截尾Truncation	无None	有Yes	中Medium
30	卵形Ovoid	不对称Asymmetry	尖Pointed	截尾Truncation	有Yes	有Yes	多Many
31	椭圆形Elliptic	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	少Few
32	卵形Ovoid	稍不对称Slightly asymmetric	尖Pointed	圆Round	有Yes	有Yes	少Few
33	圆球形Spheroid	完全对称Complete symmetry	圆Round	圆Round	无None	有Yes	多Many
34	椭圆形Elliptic	稍不对称Slightly asymmetric	圆Round	圆Round	无None	有Yes	少Few
35	卵形Ovoid	不对称Asymmetry	尖Pointed	圆Round	有Yes	有Yes	中Medium

序号 No.	形状 Shape	叶尖 Leaf apex	叶基 Leaf base	叶片弯曲形状 Blade bending shape
1	椭圆披针形Elliptical lanceolate	渐尖Acuminate	楔形Wedge	平直Flat
2	椭圆披针形Elliptical lanceolate	尖Pointed	狭楔形Narrow wedge	向上弯曲Bend upward
3	披针形Lanceolate	渐尖Acuminate	狭楔形Narrow wedge	向上弯曲Bend upward
4	长椭圆形Oblong	尖Pointed	楔形Wedge	平直Flat
5	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	扭曲Twist
6	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	向上弯曲Bend upward
7	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
8	长椭圆形Oblong	尖Pointed	阔楔形Wide wedge	向上弯曲Bend upward
9	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
10	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
11	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
12	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
13	椭圆披针形Elliptical lanceolate	尖Pointed	阔楔形Wide wedge	平直Flat
14	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	向下弯曲Bend downward
15	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
16	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
17	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
18	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
19	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
20	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
21	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
22	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
23	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
24	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	向上弯曲Bend upward
25	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
26	长椭圆形Oblong	凸尖Mucronate	阔楔形Wide wedge	平直Flat
27	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
28	椭圆披针形Elliptical lanceolate	尖Pointed	狭楔形Narrow wedge	向上弯曲Bend upward
29	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
30	披针形Lanceolate	渐尖Acuminate	狭楔形Narrow wedge	平直Flat
31	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat
32	椭圆披针形Elliptical lanceolate	渐尖Acuminate	狭楔形Narrow wedge	平直Flat
33	椭圆披针形Elliptical Lanceolate	渐尖Acuminate	楔形Wedge	扭曲Twist
34	椭圆披针形Elliptical lanceolate	渐尖Acuminate	楔形Wedge	平直Flat
35	椭圆披针形Elliptical lanceolate	尖Pointed	楔形Wedge	平直Flat

Analysis and evaluation of phenotypic character diversity of olive germplasm resources

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序号 No.	果实纵径 FVD/mm			果实横径 FTD/mm			果形指数 FSI			单果重 SFW/g			果肉厚 PT/mm			果核纵径 KVD/mm			果核横径 KTD/mm		果核指数 KSI
1	20.77± 1.47 gh			14.63± 1.01 jk			1.42± 0.08 defgh			2.75± 0.44 gh			3.36± 0.57 ghi			15.75± 1.64 efghi			6.93± 0.59 ijkl		2.27± 0.22 bcd
2	19.92± 1.29 hi			15.23± 0.89 hij			1.31± 0.07 jklmnop			2.78± 0.41 gh			3.78± 0.44 fgh			13.68± 1.14 klmn			7.38± 0.37 ghi		1.85± 0.18 jklm
3	21.92± 1.99 efg			18.25± 1.27 de			1.20± 0.06 qrst			4.40± 0.87 cd			4.97± 0.62 bcd			14.90± 1.50 ghijk			8.45± 0.72 bcd		1.76± 0.10 lmnop
4	21.96± 1.79 efg			16.21± 1.09 gh			1.35± 0.10 efghijkl			3.24± 0.51 fg			4.43± 0.57 def			15.88± 1.24 efgh			7.63± 0.38 fgh		2.08± 0.14 efgh
5	21.57± 1.88efg			17.82± 1.19 def			1.21± 0.07 qrst			3.85± 0.65 def			4.58± 0.56 cde			14.55± 1.76 ijkl			7.90± 1.21 efg		1.80± 0.22 klmn
6	24.26± 1.45 cd			17.62± 1.37 def			1.38± 0.09 efghijk			4.57± 0.71 c			4.80± 0.76 bcd			17.05± 1.44 cde			8.09± 0.56 def		2.11± 0.20 def
7	19.12± 1.37 ij			15.93± 0.97 ghi			1.20± 0.07 qrst			3.04± 0.47 g			4.81± 0.61 bcd			12.92± 1.36 mno			7.73± 0.45 efgh		1.67± 0.15 nopq
8	18.93± 1.44 ijk			14.11± 0.84 jk			1.34± 0.07 ghijklm			2.35± 0.39 hi			3.42± 0.49 ghi			14.11± 1.29 jklm			7.05± 0.48 ijkl		2.00± 0.16 fghijk
9	13.76± 0.84 qr			10.33± 0.86 qr			1.33± 0.09 hijklmn			0.99± 0.20 lm			1.79± 0.20 l			12.18± 0.78 op			6.64± 0.30 klmn		1.83± 0.10 klmno
序号 No.	果实纵径 FVD/mm			果实横径 FTD/mm			果形指数 FSI				单果重 SFW/g			果肉厚 PT/mm		果核纵径 KVD/mm			果核横径 KTD/mm		果核指数 KSI
10	15.98± 0.83 nop			11.61± 0.83 mnop			1.38± 0.07 efghijk				1.40± 0.25 jkl			2.15± 0.41 jkl		13.40± 1.01 lmno			7.09± 0.45 ijk		1.89± 0.11 ijkl
11	15.26± 0.96 opq			11.95± 0.89 mno			1.28± 0.05l mnopqr				1.36± 0.28 jklm			1.94± 0.52 l		12.97± 0.72 mno			7.68± 0.48 efgh		1.69± 0.12 mnopq
12	22.93± 1.65 de			18.66± 1.49 cde			1.23± 0.06 pqrs				4.61± 1.09 c			5.27± 0.93 bc		14.89± 1.13 ghijk			7.73± 0.47 efgh		1.93± 0.16 fghijkl
13	25.17± 1.17 bc			18.05± 1.04 def			1.39± 0.05 efghij				4.73± 0.68 c			4.97± 0.32 bcd		17.92± 1.07 bcd			8.09± 0.42 def		2.22± 0.12 cde
14	14.02± 0.84 qr			12.48± 0.69 lmn			1.12± 0.05 t				1.42± 0.22 jkl			3.07± 0.63 hi		10.05± 0.59 q			6.37± 0.35 mno		1.58± 0.07 q
15	17.59± 0.94 klm			12.49± 0.62 lmn			1.41± 0.05 efghi				1.79± 0.23 ijk			3.20± 0.34 ghi		14.33± 0.93 jkl			7.01± 0.26 ijkl		2.04± 0.12 efghi
16	21.79± 1.61 efg			17.60± 0.90 def			1.24± 0.07 opqrs				3.75± 0.51 ef			5.08± 0.34 bcd		15.16± 1.23 ghij			8.00± 0.46 def		1.89± 0.16 ijkl
17	18.02± 1.08 jklm			14.53± 0.94 jk			1.24± 0.07 nopqrs				2.29± 0.35 hi			3.88± 0.32 efg		12.69± 0.95 no			7.63± 0.49 fgh		1.66± 0.14 opq
18	25.75± 1.37 bc			16.93± 0.97 fg			1.52± 0.07 bc				4.37± 0.66 cde			4.95± 0.49 bcd		20.18± 1.14 a			7.20± 0.45 hij		2.80± 0.17 a
19	18.46± 1.24 ijkl			11.57± 0.63 nop			1.60± 0.09 b				1.52± 0.21 jkl			3.04± 0.38 hi		15.77± 1.17 efghi			6.56± 0.32l mno		2.40± 0.17 b
20	17.19± 1.18 lmn			12.16± 0.83 mno			1.41± 0.06 defghi				1.56± 0.28 jkl			3.46± 0.37 ghi		12.69± 1.08 no			6.10± 0.37 o		2.08± 0.12 efg
21	25.41± 1.53bc			18.01± 1.48def			1.41± 0.17defghi				4.88± 0.62c			4.70± 0.86cd		18.36± 1.45bc			8.62± 0.51bc		2.13± 0.19 cdef
22	18.72± 1.18 ijk			10.66± 0.61 pqr			1.76± 0.11 a				1.33± 0.18 jklm			2.74± 0.29 ijk		16.17± 0.93 efg			6.06± 0.39 o		2.67± 0.23 a
23	25.39± 2.19 bc			20.15± 1.57 b			1.26± 0.05 mnopqr				5.93± 1.21 b			5.54± 0.74 b		15.67± 1.17 fghi			8.21± 0.52 cde		1.91± 0.15 hijkl
24	21.17± 0.89 fgh			13.67± 1.26 kl			1.55± 0.13 bc				2.27± 0.31 hi			3.19± 0.73 ghi		15.94± 1.05 efgh			6.98± 0.38 ijkl		2.28± 0.14 bcd
25	15.90± 1.33 nop			11.12± 0.83 opq			1.43± 0.14 def				1.21± 0.20 klm			2.06± 0.46 kl		14.05± 1.24 jklm			6.55± 0.63 lmno		2.14± 0.24 cdef
26	26.31± 1.68 ab			19.88± 1.01 bc			1.32± 0.05 ijklmno				5.98± 0.88 b			5.04± 0.53 bcd		18.95± 1.94 ab			8.86± 0.62 ab		2.14± 0.25 cdef
27	25.84± 1.96 b			19.75± 1.36 bc			1.31± 0.09 jklmnop				5.82± 0.93 b			5.27± 0.65 bc		17.99± 1.52 bcd			8.50± 0.42 bcd		2.12± 0.15 def
28	21.75± 1.24 efg			18.78± 1.03 cd			1.16± 0.06 st				4.53± 0.68 c			4.72± 0.55 cd		14.51± 0.81 ijkl			8.62± 0.46 bc		1.68± 0.14 mnopq
29	27.57± 1.87 a			21.44± 1.27 a			1.29± 0.07 klmnopq				7.63± 1.31 a			6.33± 0.36 a		16.81± 1.25 def			9.17± 0.69 a		1.84± 0.16 klmno
30	24.90± 1.42 bc			17.46± 0.96 ef			1.43± 0.09 defg				4.39± 0.62 cd			4.89± 0.70 bcd		17.70± 1.17 bcd			7.37± 0.35 hi		2.40± 0.15 b
31	18.72± 0.8 ijk			12.84± 0.67 lm			1.46± 0.07 cde				1.93± 0.27 ij			2.86± 0.40 ij		14.67± 1.05 hijkl			7.24± 0.53 hij		2.03± 0.15 fghij
32	13.68± 0.91 r			9.77± 1.00 r			1.40± 0.13 efghi				0.76± 0.12 m			2.14± 0.31 jkl		10.64± 0.74 q			5.41± 0.33 p		1.97± 0.11 fghijk
33	15.01± 0.83 pqr			12.60± 0.68 lmn			1.19± 0.06 rst				1.48± 0.17 jkl			2.71± 0.34 ijk		10.91± 0.75 pq			6.73± 0.28 jklm		1.62± 0.11 pq
34	22.31± 0.98 ef			14.86± 0.88 ijk			1.50± 0.10 cd				3.01± 0.29 g			3.79± 0.48 fgh		16.67± 0.75 def			7.28± 0.41 hi		2.29± 0.15 bc
35	16.75± 0.74mno			12.19± 0.61 mno			1.37± 0.05 efghijk				1.40± 0.18 jkl			3.13± 0.50 ghi		12.23± 0.65 op			6.11± 0.32 no		2.00± 0.12 fghijk
序号 No.	果核重 KW/g	果肉率 PP/%			叶长 LL/mm			叶宽 LW/mm				叶形指数 LI			果实水分 FMC/%			干果含油率 OCODF/%		鲜果含油率 OCOFF/%
1	0.47± 0.09 hi	82.71± 3.66 abcdefgh			51.03± 6.09 ghijkl			10.83± 0.95 ijklmn				4.71± 0.48 defghij			71.78± 0.04 bc			27.54± 0.02 klmn		7.77± 0.01 mno
2	0.48± 0.05 hi	82.32± 3.26 abcdefgh			62.81± 4.22 abcdef			10.69± 0.94 ijklmn				5.88± 0.66 abc			81.90± 0.01 a			30.74± 0.01 fghijk		5.56± 0.01 pq
3	0.67± 0.17 cdef	84.56± 3.65 abcdef			63.71± 3.42 abcdef			9.31± 0.70 n				6.85± 0.50 a			64.70± 0.02 efgh			32.93± 0.02 cdefg		11.62± 0.01 fghi
4	0.67± 0.20 cdef	78.95± 6.42 fghi			46.93± 3.59 jkl			11.96± 0.61 efghijklm				3.92± 0.38 jk			32.79± 0.02 r			31.25± 0.01 efghij		21.00± 0.02 b
5	0.56± 0.12 efgh	85.40± 2.19 abcde			66.42± 4.97 abcd			12.68± 0.99 cdefghij				5.24± 0.49 bcdefgh			58.78± 0.03 jklmn			28.53± 0.01 jklm		11.76± 0.02 fghi
6	0.74± 0.12 cd	83.46± 3.40 abcdefgh			64.25± 5.16 abcde			13.16± 1.32 bcdefghi				4.88± 0.33 bcdefghij			60.76± 0.03 hijkl			36.41± 0.01 c		14.29± 0.01 cd
7	0.51± 0.10 ghi	83.33± 2.49 abcdefgh			47.39± 3.33 ijkl			10.91± 0.94 ijklmn				4.34± 0.38 efghijk			49.99± 0.02 p			26.68± 0.01 lmno		13.34± 0.01 cdef
8	0.44± 0.08 hij	80.90± 4.52 cdefgh			51.77± 9.66 ghijkl			12.75± 1.70 cdefghij				4.06± 0.91 hijk			68.87± 0.03 cde			32.35± 0.01 efghi		10.07± 0.01 hijkl
9	0.36± 0.04 ijkl	62.76± 7.42 n			63.02± 9.18 abcdef			13.78± 0.77 abcdefg				4.57± 1.37 defghijk			55.06± 0.01mno			18.10± 0.01 s		8.13± 0.01 lmno
10	0.43± 0.08 hijk	69.00± 3.48 lm			64.95± 3.65 abcde			15.07± 1.39 abc				4.82± 0.55 cdefghij			69.72± 0.02 cd			23.30± 0.02 opq		7.05± 0.01 nop
11	0.50± 0.06 ghi	62.12± 7.68 n			70.08± 3.20 ab			14.55± 1.16 abcd				4.31± 0.36 efghijk			45.12± 0.01 q			20.05± 0.03 qrs		4.27± 0.01 q
12	0.64± 0.10 defg	85.82± 2.24 abcd			48.64± 5.85 hijkl			9.54± 0.92 mn				5.10± 0.52 bcdefghi			56.26± 0.02 lmno			32.67± 0.01 defgh		14.29± 0.01 cd
13	0.75± 0.09 cd	83.84± 2.86 abcdefg			68.89± 5.49 abc			16.08± 1.32 a				4.28± 0.20 fghijk			68.22± 0.02 cde			34.47± 0.01 cde		10.95± 0.01 ghij
14	0.30± 0.05 jklm	78.30± 3.76 ghij			47.32± 2.01 ijkl			10.00± 0.75 klmn				4.73± 0.40 defghij			59.58± 0.02 ijklm			36.08± 0.01 cd		14.58± 0.01 cd
15	0.49± 0.05 ghi	72.65± 2.24 klm			60.50± 3.81 bcdefg			12.35± 1.06 defghijkl				4.90± 0.46 bcdefghij			57.34± 0.02 klmno			61.30± 0.03 a		26.15± 0.01 a
16	0.55± 0.09 fgh	85.15± 1.89 abcde			72.22± 7.16 a			13.20± 1.10 bcdefghi				5.47± 0.49 bcde			59.64± 0.01 ijklm			24.68± 0.01 nop		9.96± 0.01 hijkl
17	0.45± 0.07 hij	79.95± 4.60 efgh			57.36± 4.20 defghi			14.14± 1.16 abcde				4.06± 0.25 ijk			62.57± 0.02 fghij			28.89± 0.02 ijklm		10.81± 0.01 hijk
18	0.80± 0.12 bc	81.53± 2.65 cdefgh			68.55± 4.49 abc			13.94± 1.89 abcdef				4.92± 0.58 bcdefghij			59.34± 0.03 jklmn			32.44± 0.02 efghi		13.19± 0.01 cdef
19	0.42± 0.06 hijkl	72.14± 2.52 lm			62.84± 6.13 abcdef			14.16± 0.86 abcde				4.44± 0.22 defghijk			61.70± 0.07 hijk			23.20± 0.01 opq		8.89± 0.01 klmn
20	0.30± 0.05 jklm	80.52± 1.93 defgh			51.28± 3.10 ghijkl			9.88± 1.33 lmn				5.19± 0.74 bcdefgh			64.25± 0.02 efghi			26.13± 0.02 mno		9.34± 0.01 jklm
21	0.79± 0.11 bcd	83.70± 2.47 abcdefg			59.51± 3.86 cdefg			11.25± 1.09 hijklmn				5.29± 0.66 bcdefg			53.77± 0.02 op			29.15± 0.01 hijklm		13.48± 0.01 cdef
22	0.39± 0.04 ijkl	70.32± 4.60 lm			60.37± 4.98 bcdefg			14.03± 1.11 abcdef				4.30± 0.32 fghijk			57.53± 0.02 klmno			24.74± 0.01 nop		10.51± 0.01 hijk
23	0.71± 0.12 cde	87.50± 3.64 a			63.66± 3.36 abcdef			11.57± 1.24 fghijklmn				5.50± 0.53 bcd			85.56± 0.03 a			27.51± 0.01 klmn		3.97± 0.01 q
24	0.49± 0.08 ghi	77.96± 6.05 hijk			56.49± 4.91 defghij			12.97± 0.85 bcdefghi				4.36± 0.34 efghijk			54.69± 0.02 nop			30.45± 0.01 fghijk		13.80± 0.01 cde
25	0.39± 0.05 ijkl	67.58± 4.50 mn			60.99± 4.45 bcdefg			11.65± 0.7 4efghijklmn				5.24± 0.54 bcdefgh			58.08± 0.02 jklmno			23.46± 0.02 opq		9.84± 0.02 ijkl
26	1.00± 0.12 a	83.20± 1.58 abcdefgh			43.46± 1.26 l			12.43± 0.97 defghijk				3.52± 0.35 k			67.08± 0.02 cdef			20.44± 0.02 qrs		6.72± 0.01 op
序号 No.	果核重 KW/g		果肉率 PP/%			叶长 LL/mm			叶宽 LW/mm			叶形指数 LI			果实水分 FMC/%		干果含油率 OCODF/%			鲜果含油率 OCOFF/%
27	0.79± 0.10bcd		86.27± 1.77 abc			58.39± 4.88 defgh			10.80± 1.33 ijklmn			5.41± 0.52 bcdef			62.06± 0.01 ghijk		33.86± 0.02 cdef			12.85± 0.01 defg
28	0.73± 0.09 cd		83.83± 1.34 abcdefg			54.09± 3.11 fghijk			12.73± 0.86 cdefghij			4.25± 0.22 ghijk			62.48± 0.02 fghij		18.59± 0.01 rs			6.97± 0.01 nop
29	0.93± 0.27 ab		87.77± 3.18 a			60.04± 4.50 bcdefg			11.37± 0.79 ghijklmn			5.28± 0.51 bcdefg			74.96± 0.01 b		29.90± 0.01 ghijkl			7.49± 0.01 mnop
30	0.55± 0.07 fgh		87.30± 2.11 ab			55.58± 4.41 efghijk			9.23± 1.00 n			6.02± 0.62 ab			68.53± 0.02 cde		29.64± 0.02 ghijklm			9.33± 0.01 jklm
31	0.51± 0.10 ghi		73.86± 3.14 ijkl			68.53± 4.95 abc			15.30± 1.41 ab			4.48± 0.33 defghijk			70.42± 0.02 bcd		22.14± 0.02 pqr			6.55± 0.01 op
32	0.20± 0.04 m		73.09± 5.42 jklm			46.27± 3.86 kl			10.32± 0.85 jklmn			4.49± 0.70 defghijk			66.81± 0.01 defg		31.57± 0.02 efghij			10.48± 0.02 hijk
33	0.28± 0.04 klm		81.15± 2.93 cdefgh			63.68± 5.47 abcdef			12.48± 2.13 defghijk			5.10± 0.67 bcdefghi			66.95± 0.02 def		36.07± 0.01 cd			11.92± 0.03 efgh
34	0.55± 0.06 fgh		81.67± 2.45 bcdefgh			58.36± 3.92 defgh			12.01± 1.40 efghijklm			5.24± 0.56 cdefghij			53.63± 0.02 op		32.75± 0.02 defg			15.18± 0.01 c
35	0.27± 0.03 lm		80.84± 2.07 cdefgh			59.23± 4.99 cdefg			13.58± 1.27 abcdefgh			4.36± 0.16 efghijk			53.87± 0.02 op		40.03± 0.02 b			15.04± 0.01 c

性状Trait	最大值Max	最小值Min	平均值Mean	极差Range	标准差Standard deviation	变异系数CV/%
果实纵径FVD/mm	27.57	13.68	20.52	13.88	3.94	19.69
果实横径FTD/mm	21.44	9.77	15.29	11.67	3.18	21.46
果形指数FSI	1.76	1.12	1.35	0.63	0.13	9.82
单果重SFW/g	7.63	0.76	3.17	6.87	1.71	55.87
果肉厚PT/mm	6.33	1.79	3.94	4.54	1.15	30.75
果核纵径KVD/mm	20.18	10.05	14.95	10.13	2.39	16.06
果核横径KTD/mm	9.17	5.41	7.42	3.77	0.87	12.02
果核指数KSI	2.80	1.58	2.02	1.22	0.28	14.02
果核重KW/g	1.00	0.20	0.55	0.80	0.19	35.30
果肉率PP/%	87.77	62.12	79.47	25.64	6.86	8.64
叶长LL/mm	72.22	43.46	58.86	28.76	7.60	12.91
叶宽LW/mm	16.08	9.23	12.29	6.85	1.79	14.56
叶形指数LI	6.85	3.50	4.84	3.35	0.67	13.76
果实水分FMC/%	85.56	32.79	61.85	52.77	9.80	15.85
干果含油率OCODF/%	61.30	18.10	29.66	43.20	7.71	26.00
鲜果含油率OCOFF/%	26.15	3.97	11.06	22.18	4.43	40.02

性状	FVD	FTD	FSI	SFW	PT	KVD	KTD	KSI	KW	PP	LL	LW	LI	FMC	OCODF
Trait
FTD	0.901^**
FSI	0.057	-0.373^*
SFW	0.931^**	0.974^**	-0.247
PT	0.890^**	0.957^**	-0.289	0.933^**
KVD	0.888^**	0.636^**	0.415^*	0.703^**	0.624^**
KTD	0.765^**	0.892^**	-0.412^*	0.869^**	0.780^**	0.564^**
KSI	0.390^*	-0.021	0.870^**	0.080	0.067	0.696^**	-0.195
KW	0.901^**	0.879^**	-0.105	0.916^**	0.806^**	0.811^**	0.893^**	0.186
PP	0.696^**	0.782^**	-0.318	0.723^**	0.827^**	0.392^*	0.520^**	0.004	0.496^**
LL	0.070	0.000	0.150	0.011	-0.040	0.174	0.137	0.098	0.065	-0.157
LW	-0.150	-0.273	0.307	-0.246	-0.329	0.084	-0.043	0.148	-0.009	-0.451^**	0.568^**
LI	0.250	0.314	-0.186	0.295	0.334^*	0.095	0.204	-0.063	0.089	0.350^*	0.359^*	-0.556^**
FMC	0.182	0.211	-0.072	0.263	0.170	0.037	0.103	-0.045	0.078	0.319	0.196	-0.118	0.338^*
OCODF	0.004	-0.016	-0.005	-0.025	0.078	-0.038	-0.139	0.048	-0.086	0.220	-0.029	-0.199	0.182	-0.052
OCOFF	-0.017	-0.070	0.081	-0.107	0.052	0.030	-0.144	0.137	-0.053	0.076	-0.199	-0.154	-0.051	-0.580^**	0.781^**

性状 Trait	主成分Principal component
性状 Trait	1	2	3	4	5
果实纵径FVD	0.955	0.271	0.061	0.019	-0.055
果实横径 FTD	0.982	-0.13	-0.03	-0.097	0.0270
果形指数FSI	-0.219	0.875	0.158	0.27	-0.208
单果重SFW	0.983	-0.022	-0.049	-0.053	-0.021
果肉厚PT	0.953	-0.104	0.117	-0.029	-0.003
果核纵径KVD	0.743	0.643	0.08	0.016	-0.051
果核横径KTD	0.866	-0.103	-0.225	-0.293	0.216
果核指数KSI	0.129	0.865	0.268	0.276	-0.255
果核重KW	0.906	0.212	-0.072	-0.259	0.058
果肉率PP	0.776	-0.278	0.218	0.171	-0.116
叶长LL	0.041	0.329	-0.431	0.363	0.739
叶宽LW	-0.266	0.579	-0.421	-0.322	0.460
叶形指数LI	0.351	-0.316	0.027	0.732	0.212
果实水分FMC	0.248	-0.164	-0.459	0.626	-0.158
干果含油率	0.008	-0.117	0.768	0.246	0.417
OCODF
鲜果含油率	-0.061	0.033	0.918	-0.155	0.327
OCOFF
特征值	6.803	2.747	2.234	1.592	1.286
Eigenvalue
贡献率	42.516	17.166	13.962	9.951	8.038
Contribution rate/%
累计贡献率	42.516	59.682	73.644	83.595	91.633
Cumulative contribution rate/%

分组 Group	FVD/ mm	FTD/ mm	FSI	SFW/g	PT/ mm	KVD/ mm	KTD/ mm	KSI	KW/g	PP/%	LL/mm	LW/ mm	LI	FMC/ %	OCODF/ %	OCOFF/ %
Ⅰ	16.69	11.44	1.46	1.39	2.37	14.17	6.83	2.07	0.55	68.26	64.40	14.08	4.59	59.66	22.14	7.89
Ⅱ	21.34	16.13	1.32	3.58	4.27	15.15	7.55	2.01	0.57	82.59	57.31	11.88	4.82	62.07	30.51	11.47
Ⅲ	22.65	17.69	1.28	4.36	4.66	14.68	7.79	1.88	0.60	84.91	63.24	11.13	5.68	83.73	29.13	4.77
Ⅳ	21.96	16.21	1.25	3.24	4.43	15.88	7.62	2.08	0.67	78.95	45.92	11.96	3.92	32.79	31.25	21.00
Ⅴ	17.59	12.49	1.41	1.79	3.20	14.33	7.01	2.04	0.49	72.65	60.50	12.35	4.90	57.34	61.30	26.15

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