枇杷生物活性物质积累机制的研究进展

doi:10.3969/j.issn.1004-1524.20250707

浙江农业学报 ›› 2026, Vol. 38 ›› Issue (8): 1706-1716.DOI: 10.3969/j.issn.1004-1524.20250707

枇杷生物活性物质积累机制的研究进展

汪彬君¹^,²(), 葛航², 李晓颖², 谢宇迪³, 陈俊伟², 高永彬¹^,^*(), 徐红霞²^,^*()

¹ 浙江农林大学园艺科学学院, 浙江杭州 311300
² 浙江省农业科学院园艺研究所, 浙江杭州 310021
³ 河北科技师范学院园艺科技学院, 河北秦皇岛 066600

收稿日期:2025-11-09 出版日期:2026-08-25 发布日期:2026-09-14
作者简介:汪彬君,研究方向为果树育种与栽培。E-mail:17857323821@139.com
通讯作者: *高永彬,E-mail:gaoybin@zafu.edu.cn;徐红霞,E-mail:xuhongxia@zaas.ac.cn
基金资助:
浙江省“三农九方”科技协作项目(2025SNJF064);兰溪市特色水果团队特派员项目;浙江省“十四五”果品新品种选育专项(2021C02066-3);浙江省自然科学基金(LTGN24C150006)

Research progress on the accumulation mechanism of bioactive compounds in Eriobotrya japonica

WANG Binjun¹^,²(), GE Hang², LI Xiaoying², XIE Yudi³, CHEN Junwei², GAO Yongbin¹^,^*(), XU Hongxia²^,^*()

¹ College of Horticultural Science, Zhejiang A&F University, Hangzhou 311300, China
² Institute of Horticulture, Zhejiang Academy of Agricultural Sciences, Hangzhou 310021, China
³ College of Horticulture Science & Technology, Hebei Normal University of Science & Technology, Qinhuangdao 066600, Hebei, China

Received:2025-11-09 Published:2026-08-25 Online:2026-09-14

摘要/Abstract

摘要：

枇杷(Eriobotrya japonica)是一种营养与药用价值兼备的蔷薇科植物,其果实、叶片、种子和花中富含多种生物活性物质,如黄酮类、酚酸类、三萜酸类、挥发油和多糖,这些物质赋予了枇杷抗氧化、抗炎、抗肿瘤、降血糖和护肺等多种药理活性。生物活性物质的含量和组成是决定枇杷品质和功效的关键,而它们的积累受到复杂的内部机制和外部环境的共同调控。本文系统综述了近年来枇杷主要生物活性物质的种类与功能研究进展,重点阐述了其生物合成途径、关键酶基因调控、特异性积累规律,以及分子调控机制的研究进展,指出当前研究中存在的机制不明、多因子交互作用研究薄弱等问题,并展望了多组学整合分析、基因编辑应用和产学研转化等未来发展方向,以期为枇杷活性成分的高效利用和产业增值提供理论支撑。

关键词: 枇杷, 酚类, 黄酮, 三萜酸, 积累机制

Abstract:

Loquat (Eriobotrya japonica) is a plant of the Rosaceae family valued for both its nutritional and medicinal properties. Its fruits, leaves, seeds and flowers are rich in various bioactive compounds, such as flavonoids, phenolic acids, triterpenic acids, essential oils and polysaccharides. These substances confer multiple pharmacological activities to loquat, including antioxidant, anti-inflammatory, antitumor, hypoglycemic, and lung-protective effects. The content and composition of these bioactive compounds are crucial determinants of loquat quality and efficacy, and their accumulation is jointly regulated by complex internal mechanisms and external environmental factors. This systematic review summarizes recent research progress on the types and functions of major bioactive compounds in loquat, focusing on advances in their biosynthetic pathways, key enzyme gene regulation, specific accumulation patterns, and molecular regulatory mechanisms. It highlights existing gaps in understanding, such as unclear mechanisms and insufficient investigation of multifactorial interactions, while outlining future directions including multi-omics integration analysis, gene editing applications, and industry-academia-research collaboration. These efforts aim to provide theoretical support for the efficient utilization of loquat bioactive components and industrial value enhancement.

Key words: loquat, phenolic, flavonoid, triterpenic acid, accumulation mechanism

中图分类号:

S667.3

汪彬君, 葛航, 李晓颖, 谢宇迪, 陈俊伟, 高永彬, 徐红霞. 枇杷生物活性物质积累机制的研究进展[J]. 浙江农业学报, 2026, 38(8): 1706-1716.

WANG Binjun, GE Hang, LI Xiaoying, XIE Yudi, CHEN Junwei, GAO Yongbin, XU Hongxia. Research progress on the accumulation mechanism of bioactive compounds in Eriobotrya japonica[J]. Acta Agriculturae Zhejiangensis, 2026, 38(8): 1706-1716.

参考文献 74

[1]	林顺权. 栽培枇杷国际传播史考[J]. 果树学报, 2020, 37(2): 272-277.
	Lin S Q. Verification of the international introduction history of cultivated loquats[J]. Journal of Fruit Science, 2020, 37(2): 272-277.
[2]	林国荣, 沈高扬. 枇杷核仁的营养成分分析[J]. 食品科技, 2007, 32(8): 276-277.
	Lin G R, Shen G Y. Analyzed nutrition composition of the Eriobotrya japonia nut[J]. Food Science and Technology, 2007, 32(8): 276-277.
[3]	李鹰, 唐晓云, 赵启超, 等. 枇杷核提取物对机体免疫功能影响的探讨[J]. 中医药信息, 1999, 16(1): 51-52.
	Li Y, Tang X Y, Zhao Q C, et al. Effect of loquat seed extract on immune function[J]. Information on Traditional Chinese Medicine, 1999, 16(1): 51-52.
[4]	Da Silva A P G, Sganzerla W G, John O D, et al. A comprehensive review of the classification, sources, biosynthesis, and biological properties of hydroxybenzoic and hydroxycinnamic acids[J]. Phytochemistry Reviews, 2025, 24(2): 1061-1090.
[5]	陈志杰, 吴嘉琪, 马燕, 等. 植物食品原料中酚酸的生物合成与调控及其生物活性研究进展[J]. 食品科学, 2018, 39(7): 321-328.
	Chen Z J, Wu J Q, Ma Y, et al. Biosynthesis, metabolic regulation and bioactivity of phenolic acids in plant food materials[J]. Food Science, 2018, 39(7): 321-328.
[6]	张文娜, 李鲜, 孙崇德, 等. 枇杷酚类物质及其生物活性研究进展[J]. 食品与药品, 2015, 17(2): 123-128.
	Zhang W N, Li X, Sun C D, et al. Phenolic compounds in loquat and their bioactivities[J]. Food and Drug, 2015, 17(2): 123-128.
[7]	Zhang K, Ma Q L, Wang Y, et al. Transcriptome and biochemical analyses reveal phenolic compounds-mediated flavor differences in loquat (Eriobotrya japonica Lindl.) cultivars Chunhua No.1 and Dawuxing[J]. Food Chemistry: X, 2024, 21: 101145.
[8]	王增斌, 宋雪, 王洋, 等. 基于UHPLC-MS技术的枇杷叶次生代谢物周年变化研究[J]. 西北农林科技大学学报(自然科学版), 2023, 51(2): 43-52.
	Wang Z B, Song X, Wang Y, et al. Annual variation of secondary metabolites in loquat leaves based on UHPLC-MS technology[J]. Journal of Northwest A&F University(Natural Science Edition), 2023, 51(2): 43-52.
[9]	王建超. 枇杷叶多酚提取、纯化及其抗氧化、抑菌活性研究[D]. 福州: 福建农林大学, 2015.
	Wang J C. Study on extraction and purification process, antioxidant and antibacterial activity of polyphenols from loquat leaves[D]. Fuzhou: Fujian Agriculture and Forestry University, 2015.
[10]	Xu H X, Li X Y, Chen J W. Comparison of phenolic compound contents and antioxidant capacities of loquat (Eriobotrya japonica Lindl.) fruits[J]. Food Science and Biotechnology, 2014, 23(6): 2013-2020.
[11]	周立稳, 彭效明, 居瑞军, 等. 不同溶剂对枇杷果提取物抗氧化活性和多酚稳定性的影响[J]. 食品与生物技术学报, 2024, 43(9): 107-115.
	Zhou L W, Peng X M, Ju R J, et al. Effect of different solvents on the antioxidant activity and polyphenol stability of loquat (Eriobotrya japonica Lindl.) fruit extract[J]. Journal of Food Science and Biotechnology, 2024, 43(9): 107-115.
[12]	王萍, 王宇鹤, 李辉, 等. 枇杷核不同极性萃取物总黄酮、总多酚含量与其抗氧化活性的相关性[J]. 化学试剂, 2020, 42(9): 1067-1072.
	Wang P, Wang Y H, Li H, et al. Correlation between the content total flavonoids and total polyphenols in loquat nuclear with different extracts and their antioxidant activities[J]. Chemical Reagents, 2020, 42(9): 1067-1072.
[13]	周明国. 植物化学[M]. 2版. 北京: 科学出版社, 2012: 141.
[14]	Shen N, Wang T F, Gan Q, et al. Plant flavonoids: classification, distribution, biosynthesis, and antioxidant activity[J]. Food Chemistry, 2022, 383: 132531.
[15]	邵长芬, 李得发. 地枇杷的活性成分生物学功能及在动物生产中的应用[J]. 农业与技术, 2023, 43(8): 73-76.
	Shao C F, Li D F. Biological function of active components of loquat and its application in animal production[J]. Agriculture and Technology, 2023, 43(8): 73-76.
[16]	李佳美, 徐伟, 张雪, 等. 基于HPLC-MS/MS对枇杷叶水提组分分析及萜类物质的结构鉴定[J]. 食品工业科技, 2022, 43(1): 295-303.
	Li J M, Xu W, Zhang X, et al. HPLC-MS/MS analysis of water-extracted components of loquat leaves and structural identification of terpenoids[J]. Science and Technology of Food Industry, 2022, 43(1): 295-303.
[17]	玉澜, 黄宏清, 吴承武, 等. 超声波辅助双水相提取枇杷叶中的黄酮及其抗氧化研究[J]. 饲料研究, 2022, 45(1): 67-71.
	Yu L, Huang H Q, Wu C W, et al. Ultrasonic-assisted two-phase extraction of flavonoids from loquat leaves and their antioxidant activity[J]. Feed Research, 2022, 45(1): 67-71.
[18]	姜帆, 周丹蓉, 高慧颖, 等. 枇杷种质花中黄酮含量分析与评价[J]. 植物遗传资源学报, 2016, 17(6): 1031-1035.
	Jiang F, Zhou D R, Gao H Y, et al. Analysis and evaluation of flavonoid content in flowers of loquat [Eriobotrya japonica(Thunb.) Lindl.] germplasm resources[J]. Journal of Plant Genetic Resources, 2016, 17(6): 1031-1035.
[19]	毕云华, 黄佳婷, 章鸿彬, 等. 枇杷花抗炎活性物质的制备及成分分析[J]. 食品与发酵工业, 2024, 50(12): 178-184.
	Bi Y H, Huang J T, Zhang H B, et al. Preparation and analysis of anti-inflammatory substances from loquat flowers[J]. Food and Fermentation Industries, 2024, 50(12): 178-184.
[20]	Zeng T, Chen Y, Jian Y X, et al. Chemotaxonomic investigation of plant terpenoids with an established database (TeroMOL)[J]. New Phytologist, 2022, 235(2): 662-673.
[21]	牟玉兰, 闫浩, 龚黎黎. 萜类化合物的研究概况[J]. 化工管理, 2018(11): 12-13.
	Mou Y L, Yan H, Gong L L. Research survey of terpenoids[J]. Chemical Enterprise Management, 2018(11): 12-13.
[22]	Zhang C B, Li M, Zhao G R, et al. Alpha-terpineol production from an engineered Saccharomyces cerevisiae cell factory[J]. Microbial Cell Factories, 2019, 18(1): 160.
[23]	陈剑, 赵磊, 黄晓杰, 等. 枇杷叶6种三萜酸的人肠道菌群生物转化及其抑制α-葡萄糖苷酶活性研究[J]. 时珍国医国药, 2022, 33(2): 270-272.
	Chen J, Zhao L, Huang X J, et al. Biotransformation of six triterpene acids from Eriobotrya japonica leaves by human intestinal bacteria and their relation to in vitro α-glucosidase inhibitory activity[J]. Lishizhen Medicine and Materia Medica Research, 2022, 33(2): 270-272.
[24]	陈秀萍, 苏文炳, 蒋际谋, 等. 两种提取法的枇杷果肉萜类化合物组分含量差异分析[J]. 东南园艺, 2022, 10(2): 99-106.
	Chen X P, Su W B, Jiang J M, et al. Difference analysis on the compounds and contents of terpenoids of loquat fresh between two extraction methods[J]. Fujian Fruits, 2022, 10(2): 99-106.
[25]	Su W B, Jing Y, Lin S K, et al. Polyploidy underlies co-option and diversification of biosynthetic triterpene pathways in the apple tribe[J]. Proceedings of the National Academy of Sciences of the United States of America, 2021, 118(20): e2101767118.
[26]	刘小洪, 孙海艳, 陈薇薇, 等. 枇杷果实类胡萝卜素研究进展[J]. 果树学报, 2016, 33(7): 874-881.
	Liu X H, Sun H Y, Chen W W, et al. Advances in studies on carotenoids in loquat fruit[J]. Journal of Fruit Science, 2016, 33(7): 874-881.
[27]	韩永. 两个枇杷品种成熟期果肉生理生化与蛋白质组分析[D]. 福州: 福建农林大学, 2014.
	Han Y. Analysis of physiology, biochemistry and proteomic of two loquat cultivar pulp at the mature stage[D]. Fuzhou: Fujian Agriculture and Forestry University, 2014.
[28]	杨婧娀, 王佐民, 赵云财. 枇杷叶发酵酒[J]. 酿酒, 2018, 45(2): 97-98.
	Yang J S, Wang Z M, Zhao Y C. Loquat leaf fermented wine[J]. Liquor Making, 2018, 45(2): 97-98.
[29]	梁玉文, 冯学梅, 李阿波, 等. 宁夏日光温室枇杷果实品质分析研究试验[J]. 农业科技通讯, 2013(9): 103-104.
	Liang Y W, Feng X M, Li A B, et al. Study on fruit quality analysis of loquat in Ningxia solar greenhouse[J]. Bulletin of Agricultural Science and Technology, 2013(9): 103-104.
[30]	魏秀清, 邓朝军, 章希娟, 等. 枇杷种质资源果实维生素C与总酸含量分析[J]. 福建果树, 2009(3): 30-33.
	Wei X Q, Deng C J, Zhang X J, et al. Analysis of vitamin C and total acid contents of loquat germplasm resources[J]. Fujian Fruits, 2009(3): 30-33.
[31]	刘丽丽, 刘玉垠, 王杰, 等. 枇杷功能成分及生物活性研究进展[J]. 食品科学, 2020, 41(5): 306-314.
	Liu L L, Liu Y Y, Wang J, et al. A review of functional components and bioactivities in loquat (Eriobotrya japonica Lindl.)[J]. Food Science, 2020, 41(5): 306-314.
[32]	宋宁, 刘语, 刘铮铮, 等. GC-MS方法分析新鲜枇杷叶挥发油及β-葡萄糖苷酶对枇杷叶的增香作用[J]. 中国林副特产, 2019(6): 5-7.
	Song N, Liu Y, Liu Z Z, et al. Analysis of essential oil by GC-MS and the aromatization of β-glucanase on fresh Eriobotrya japonica leaves[J]. Forest By-Product and Speciality in China, 2019(6): 5-7.
[33]	芦艳, 鲁周民, 樊美丽, 等. 枇杷花开放过程挥发性物质变化研究[J]. 园艺学报, 2013, 40(11): 2245-2254.
	Lu Y, Lu Z M, Fan M L, et al. Changes of volatile components during different stages of loquat flowers[J]. Acta Horticulturae Sinica, 2013, 40(11): 2245-2254.
[34]	黄佳瑜, 薛岩伟, 王菲, 等. ‘白玉’枇杷花不同花期挥发性物质分析[J]. 食品安全质量检测学报, 2022, 13(22): 7287-7296.
	Huang J Y, Xue Y W, Wang F, et al. Analysis of volatile components in different flowering stages of ‘Baiyu’ Eriobotrya japonica flowers[J]. Journal of Food Safety and Quality, 2022, 13(22): 7287-7296.
[35]	李长虹, 秦小梅, 张璐璐, 等. 枇杷核挥发油化学成分及体外抗氧化活性研究[J]. 华中师范大学学报(自然科学版), 2014, 48(1): 58-61.
	Li C H, Qin X M, Zhang L L, et al. Study on chemical constituents and antioxidant activitise in vitro of volatile oil from loquat seed[J]. Journal of Huazhong Normal University(Natural Sciences), 2014, 48(1): 58-61.
[36]	倪敏, 凌雪峰, 王丽娟, 等. 枇杷果实挥发油中的化学成分[J]. 光谱实验室, 2013, 30(4): 1856-1858.
	Ni M, Ling X F, Wang L J, et al. Chemical constituents of volatile oil from fruit of Eriobotrya japonica[J]. Chinese Journal of Spectroscopy Laboratory, 2013, 30(4): 1856-1858.
[37]	林素英, 林毅鑫, 林清华, 等. 不同品种枇杷叶挥发油成分分析[J]. 莆田学院学报, 2020, 27(2): 71-76.
	Lin S Y, Lin Y X, Lin Q H, et al. Analysis of essential oil of loquat leaves of different varieties[J]. Journal of Putian University, 2020, 27(2): 71-76.
[38]	Gao Y C, Liu Z C, Dai S S, et al. Modification, structure identification, and biological activities of phosphorylated polysaccharides: a review[J]. Starch-Stärke, 2024, 76(7/8): 2300223.
[39]	俞一冰, 刘玉红, 左言, 等. 枇杷叶多糖的提取与纯化、结构特征及生物活性研究进展[J]. 食品与发酵科技, 2025, 61(2): 105-111.
	Yu Y B, Liu Y H, Zuo Y, et al. Research progress on extraction, purification, structural characteristics and biological activities of polysaccharides from leaves of Eriobotrya japonica[J]. Food and Fermentation Science & Technology, 2025, 61(2): 105-111.
[40]	解军. 酿酒枇杷渣中枇杷多糖的提取分离及其活性研究[D]. 福州: 福建农林大学, 2012.
	Xie J. A study about the extraction and activity of loquat polysaccharide in Saccharomyces loquat slag[D]. Fuzhou: Fujian Agriculture and Forestry University, 2012.
[41]	Zhu Q X, Li X Y, Ge H, et al. Genetic tendency analysis and comprehensive antioxidant activity evaluation of leaves and flowers of loquat F1 generation[J]. Current Issues in Molecular Biology, 2025, 47(1): 58.
[42]	林素英, 谢文燕, 何松涛, 等. 不同品种枇杷果实酚类物质及其抗氧化活性分析[J]. 食品工业科技, 2016, 37(18): 149-152.
	Lin S Y, Xie W Y, He S T, et al. Phenolic contents and antioxidant activity in the fruit of different loquat cultivars[J]. Science and Technology of Food Industry, 2016, 37(18): 149-152.
[43]	Ferreres F, Gomes D, Valentão P, et al. Improved loquat (Eriobotrya japonica Lindl.) cultivars: Variation of phenolics and antioxidative potential[J]. Food Chemistry, 2009, 114(3): 1019-1027.
[44]	吕寒, 习超鹏, 陈剑, 等. 不同枇杷叶药材中三萜和黄酮类成分含量的比较[J]. 中成药, 2009, 31(1): 89-92.
	LYU H, Xi C P, Chen J, et al. Comparison of triterpenoid and flavonoid content between the adult leaves and the defoliations of Eriobotrya japonica(Thunb.) Lindl[J]. Chinese Traditional Patent Medicine, 2009, 31(1): 89-92.
[45]	Zhou C H, Li X, Xu C J, et al. Hydrophilic and lipophilic antioxidant activity of loquat fruits[J]. Journal of Food Biochemistry, 2012, 36(5): 621-626.
[46]	Fu X M, Kong W B, Peng G, et al. Plastid structure and carotenogenic gene expression in red- and white-fleshed loquat (Eriobotrya japonica) fruits[J]. Journal of Experimental Botany, 2012, 63(1): 341-354.
[47]	李惠华, 徐剑, 刘小英, 等. 鲨烯环氧酶基因(SQEs)的克隆及其在温度胁迫下与枇杷悬浮培养细胞中熊果酸(UA)含量的相关性[J]. 农业生物技术学报, 2015, 23(4): 481-491.
	Li H H, Xu J, Liu X Y, et al. Cloning of squalene epoxidase genes (SQEs) and correlation analysis between their expression and ursolic acid (UA) content in suspension cells of loquat (Eriobotrya japonica L.) under temperature stress[J]. Journal of Agricultural Biotechnology, 2015, 23(4): 481-491.
[48]	ZHANG J, GE H, ZANG C, et al. EjODO1, a MYB transcription factor, regulating lignin biosynthesis in developing loquat (Eriobotrya japonica) fruit[J]. Frontiers in Plant Science, 2016, 7: 1360.
[49]	胡路艳. 套袋影响枇杷果实品质及转录因子EjHY5调控类胡萝卜素积累的研究[D]. 重庆: 西南大学, 2024.
	Hu L Y. Study on bagging effect on Eriobotry japonica fruit quality and regulation of carotenoid accumulation by EjHY5 transcription factor[D]. Chongqing: Southwest University, 2024.
[50]	陈秀萍, 苏文炳, 蒋际谋, 等. 枇杷不同组织中主要三萜酸含量多样性分析[J]. 果树学报, 2022, 39(4): 621-630.
	Chen X P, Su W B, Jiang J M, et al. Diversity assessment on triterpene metabolites in different tissues of ten types of loquat germplasms[J]. Journal of Fruit Science, 2022, 39(4): 621-630.
[51]	朱启轩, 李晓颖, 葛航, 等. 宁海白和大房枇杷F1代叶、花和果抗氧化活性分析与评价[J]. 果树学报, 2024, 41(9): 1716-1730.
	Zhu Q X, Li X Y, Ge H, et al. Analysis and evaluation of antioxidant activity of leaves, flowers and fruits of loquat F1 generation of Ninghaibai and Oobusa[J]. Journal of Fruit Science, 2024, 41(9): 1716-1730.
[52]	吴媛琳. 枇杷不同部位主要有效成分及抗氧化活性的比较[D]. 杨凌: 西北农林科技大学, 2014.
	Wu Y L. The main active ingredient content and the antioxidant activity of different parts of Eriobotrya japonica[D]. Yangling: Northwest A & F University, 2014.
[53]	张鑫楠, 刘艳, 李静, 等. 4个枇杷品种果肉和种子中主要功能成分差异分析[J]. 中国果树, 2025(1): 90-95.
	Zhang X N, Liu Y, Li J, et al. Difference analysis of functional components in flesh and seeds of four loquat cultivated varieties[J]. China Fruits, 2025(1): 90-95.
[54]	曹红云. 枇杷根、叶主要药效成分含量及活性比较研究[D]. 福州: 福建农林大学, 2012.
	Cao H Y. Comparison on the main medicinal components and activities of loquat root and loquat leaf[D]. Fuzhou: Fujian Agriculture and Forestry University, 2012.
[55]	张巧. 枇杷果皮的成分研究及其饮料开发[D]. 南京: 南京农业大学, 2016.
	Zhang Q. Study on composition of loquat peel and development of loquat peel beverage[D]. Nanjing: Nanjing Agricultural University, 2016.
[56]	Azuma A, Yakushiji H, Koshita Y, et al. Flavonoid biosynthesis-related genes in grape skin are differentially regulated by temperature and light conditions[J]. Planta, 2012, 236(4): 1067-1080.
[57]	胡明玥, 赵杰, 曹征宇, 等. 不同套袋处理对‘火炬’枇杷果实基础性状与品质的影响[J]. 上海农业科技, 2025(3): 96-98.
	Hu M Y, Zhao J, Cao Z Y, et al. Effects of different bagging treatments on basic characters and quality of ‘Torch’ loquat fruit[J]. Shanghai Agricultural Science and Technology, 2025(3): 96-98.
[58]	Xu H X, Chen J W, Xie M. Effect of different light transmittance paper bags on fruit quality and antioxidant capacity in loquat[J]. Journal of the Science of Food and Agriculture, 2010, 90(11): 1783-1788.
[59]	Zhang K, Chen M M, Peng X M, et al. Pre-harvest application of forchlorfenuron enhances fruit quality and bioactive substance accumulation in Chunhua loquat[J]. Food Chemistry, 2025, 473: 143059.
[60]	Yang C Q, Fang X, Wu X M, et al. Transcriptional regulation of plant secondary metabolism[J]. Journal of Integrative Plant Biology, 2012, 54(10): 703-712.
[61]	Liu W J, Wang Y C, Sun J J, et al. MdMYBDL1 employed by MdHY5 increases anthocyanin accumulation via repression of MdMYB16/308 in apple[J]. Plant Science, 2019, 283: 32-40.
[62]	Wang Y C, Liu W J, Jiang H Y, et al. The R2R3-MYB transcription factor MdMYB24-like is involved in methyl jasmonate-induced anthocyanin biosynthesis in apple[J]. Plant Physiology and Biochemistry, 2019, 139: 273-282.
[63]	Jia N, Liu J Q, Sun Y F, et al. Citrus sinensis MYB transcription factors CsMYB330 and CsMYB308 regulate fruit juice sac lignification through fine-tuning expression of the Cs4CL1 gene[J]. Plant Science, 2018, 277: 334-343.
[64]	Liu M Y, Zhang Z, Xu Z X, et al. Overexpression of SlMYB75 enhances resistance to Botrytis cinerea and prolongs fruit storage life in tomato[J]. Plant Cell Reports, 2021, 40(1): 43-58.
[65]	Wang X C, Wu J, Guan M L, et al. Arabidopsis MYB4 plays dual roles in flavonoid biosynthesis[J]. The Plant Journal, 2020, 101(3): 637-652.
[66]	Premathilake A T, Ni J B, Bai S L, et al. R2R3-MYB transcription factor PpMYB17 positively regulates flavonoid biosynthesis in pear fruit[J]. Planta, 2020, 252(4): 59.
[67]	Qian J F, You W W, Wang Y C, et al. Transcription factor EjMYBF1 positively regulates flavonol biosynthesis in loquat (Eriobotrya japonica)[J]. Plant Physiology and Biochemistry, 2025, 229: 110633.
[68]	穆红梅, 杜秀菊, 张秀省, 等. 植物MYB转录因子调控苯丙烷类生物合成研究[J]. 北方园艺, 2015(24): 171-174.
	Mu H M, Du X J, Zhang X S, et al. Study on plants MYB transcription factors regulate biological synthesis of phenylpropanoid metabolism[J]. Northern Horticulture, 2015(24): 171-174.
[69]	Gangappa S N, Botto J F. The multifaceted roles of HY5 in plant growth and development[J]. Molecular Plant, 2016, 9(10): 1353-1365.
[70]	杨雪, 雒军, 王引权. 调控植物黄酮类化合物生物合成的MYB转录因子研究进展[J]. 甘肃中医药大学学报, 2018, 35(6): 77-81.
	Yang X, Luo J, Wang Y Q. MYB transcription factors involved in regulation of flavonoid biosynthesis of medicinal plants[J]. Journal of Gansu University of Chinese Medicine, 2018, 35(6): 77-81.
[71]	El Omari N, Bakrim S, Bakha M, et al. DNA methylation: its role and interaction with epigenetic modifications in cancer[J]. Progress in Microbes & Molecular Biology, 2025, 8(1): a0000458.
[72]	李惠华, 常强, 王伟, 等. 基于转录组测序的枇杷五环三萜合成途径差异基因的分析[J]. 农业生物技术学报, 2018, 26(2): 222-233.
	Li H H, Chang Q, Wang W, et al. Differential genes analysis of pentacyclic triterpenoid synthesis pathway based on loquat (Eriobotrya japonica) transcriptome sequencing[J]. Journal of Agricultural Biotechnology, 2018, 26(2): 222-233.
[73]	薛岩伟. 枇杷五环三萜提取、鉴定及合成关键基因的挖掘与鉴定[D]. 扬州: 扬州大学, 2023.
	Xue Y W. Extraction, identification, synthesis and functional analysis of key genes of pentacyclic triterpenes in loquat[D]. Yangzhou: Yangzhou University, 2023.
[74]	Liu C, Wang M B, Wang L L, et al. Extensive genetic and DNA methylation variation contribute to heterosis in triploid loquat hybrids[J]. Genome, 2018, 61(6): 437-447.

枇杷生物活性物质积累机制的研究进展

Research progress on the accumulation mechanism of bioactive compounds in Eriobotrya japonica

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